Research Article

Molecular Detection and Distribution of Giardia duodenalis and Cryptosporidium spp. Infections in Wild and Domestic Animals in Portugal

Table 2

Prevalence and molecular diversity of Giardia duodenalis and Cryptosporidium spp. reported in domestic dogs and livestock species in Europe.

PathogenHost scientific nameCountryOccurrence rate (no. pos./total no.)Detection methodLocusSpecies/genotype (n)Subgenotype (n)Reference

Domestic carnivore
Giardia duodenalisCanis lupus familiarisItaly15 (17/113)CM, PCRssu-rRNAA (2), C (11), D (1), A + C (2), C + D (1)Berrilli et al. [54]
Italy100 (21/21)PCRssu-rRNA, gdh, bgA (6), C (1), D (13), A + D (1)A13 (5), A83 (1), D13 (7), D23 (4), D1 + D23 (2)Lalle et al. [55]
Italy19 (20/101)CM, PCRtpiA (3), C (17)A13 (3), C13 (13), C23 (4)Papini et al. [56]
Italy21 (26/127)CM, PCRssu-rRNAA (8), C (14), D (4)Scaramozzino et al. [57]
Italy64 (9/14)CM, PCRbgA (9)A13 (9)Marangi et al. [58]
Italy58 (165/285)CM, PCRssu-rRNA, bgB (1), C (49), D (29)B13 (1)Simonato et al. [59]
Italy29 (204/705)CM, PCRssu-rRNA, bgB (1), C (9), D (11)NASimonato et al. [60]
Italy21 (56/262)CM, PCRssu-rRNAC (6), D (19)Liberato et al. [61]
Italy41 (69/168)CM, PCRssu-rRNA, bg, tpiA (16), B (6), C (2), A + B (1)AII (5)Agresti et al [62]
The Netherlands100 (2/2)PCR1ssu-rRNA, gdhD (2)van der Giessen et al. [42]
The Netherlands15 (14/92)CM, PCRssu-rRNAA (1), C (7), D (3), C + D (1)Overgaauw et al. [63]
The Netherlands86 (493/573)CM, PCRssu-rRNANAUiterwijk et al. [64]
Hungary59 (110/187)CM, PCRssu-rRNAC (5), D (9), C + D (1)Szénási et al. [65]
Germany100 (92/92)PCR1ssu-rRNAC (33), D (50), C + D (8), A + D (1)Barutzki et al. [66]
Germany92 (55/60)CM, PCRssu-rRNA, gdhA (33), C (5), D (2), A + C (15)AI (14)Leonhard et al. [67]
Germany6 (5/81)PCRgdhA (5)NASotiriadou et al. [68]
Germany95 (123/130)CM, PCRssu-rRNA, gdh, bgA (24), B (8), C (52), D (69), F (6)A13 (3), A23 (1), A53 (4), A1/A53 (2), BIII (3)Pallant et al. [69]
Germany42 (13/31)CM, PCRssu-rRNA, gdh, bg, tpiA (2), C (1), D (2), A + B (1)AI (2)Rehbein et al. [70]
Germany31 (115/376)CM, PCRssu-rRNA, gdh, tpi, tpi-D2A (2), C (42), D (52), E (2), F (2)NASommer et al. [71]
Germany29 (112/386)CM, PCRssu-rRNA, gdh, bgA (9), C (43), D (40), C + D (5), D + A (4), C + A (3)AI (9)Murnik et al. [72]
Belgium23 (270/1159)CM, PCRbgA (40), B (4), C (26), D (49)A23 (2), A33 (36), D13 (16), D23 (25)Claerebout et al. [73]
United Kingdom10 (87/878)CM, PCRssu-rRNA, bgA (1), C (10), D (29), C + D (1)A33 (1), D13 (2), D23 (8)Upjohn et al. [74]
Poland2 (3/148)CM, PCRbgC (1), D (1)Solarczyk and Majewska [75]
Poland29 (31/108)CM, PCRtpiC (2)Bajer et al. [76]
Poland19 (7/36)CM, PCRbgA (1), E (1)NAStojecki et al. [77]
Poland6 (13/217)PCRbgC (10), D (3)Piekara-Stępińska et al. [78]
Spain16 (96/604)CM, PCRgdh, bgA (3), B (22), C (1), D (7), A + B (3), A + C (3), A + E (1), B + C (3), B + D (11), B + E (4), C + D (1), C + E (1), B + C + D (3)AI (10), BIII (21), BIV (25)Dado et al. [79]
Spain37 (64/169)CM, PCRgdh, bg, tpiC (1), D (4), C + D (1)Ortuño et al. [80]
Spain29 (16/55)CM, PCRssu-rRNA, gdh, bgC (3)De Lucio et al. [81]
Spain37 (127/348)CM, PCRssu-rRNA, gdh, bgA (5), B (8), C (2), D (13), A + B (4), A + D (1), A + B + D (2)AII (11), AIII (2), BIII (7), BIV (8)Adell-Aledón et al. [82]
Croatia62 (16/26)CM, PCRssu-rRNA, ITS1-5.8S-ITS2, gdh, bg, tpiC (6), D (10)Sommer et al. [83]
Macedonia33 (45/136)CM, PCRssu-rRNA, ITS1-5.8S-ITS2, gdh, bg, tpiC (7), D (7)Sommer et al. [83]
Romania36 (66/183)CM, PCRssu-rRNA, ITS1-5.8S-ITS2, gdh, bg, tpiC (8), D (8)Sommer et al. [83]
Romania100 (39/39)PCR1gdhC (8), D (29), C + D (1), E (1)Adriana et al. [27]
Serbia66 (88/134)CM, PCRssu-rRNA, ITS1-5.8S-ITS2, gdh, bg, tpiC (8), D (6)Sommer et al. [83]
Greece25 (220/879)CM, PCRssu-rRNA, gdh, bg, tpiA (2), C (45), D (27), C + D (15), A + C (2), A + D (1), B + C (1) A + C + D (2)AI (1), AII (1), BIV (1)Kostopoulou et al. [84]
Czech Republic100 (54/54)PCR1gdh, bg, tpiC (21), D (32), C + D (1)Lecová et al. [85]
Portugal17 (25/148)CM, PCRssu-rRNAB (1), C (15), D (12)Ferreira et al. [86]
Portugal34 (27/80)CM, PCRgdh, bg, tpiC (3), D (11), C + D (4)Pereira et al. [87]
Portugal24 (11/46)PCRssu-rRNA, gdh, bg, tpiD (1)This study
Cryptosporidium spp.Canis lupus familiarisCzech Republic100 (2/2)PCR1ssu-rRNA, cowpC. parvum (1), C. meleagridis (1)Hajdušek et al. [46]
Italy3 (8/240)PCRcowpC. parvum (7), C. canis (1)Giangaspero et al. [88]
Italy1 (3/285)PCRcowpC. parvum (3)Simonato et al. [59]
Italy2 (12/705)PCRssu-rRNA, cowpC. parvum (11), C. canis (1)Simonato et al. [60]
Germany1 (1/81)PCRssu-rRNAC. parvum (1)Sotiriadou et al. [68]
Germany10 (35/349)PCRssu-rRNA, gp60C. canis (33), C. parvum (2)IIa (2), XXd (5), XXe (6), XXb (2)Murnik et al. [89]
France3 (3/116)CM, PCRssu-rRNAC. canis (3)Osman et al. [90]
Greece6 (52/879)CM, PCRssu-rRNA, hsp70C. canis (2), C. scrofarum (1)Kostopoulou et al. [84]
Spain6 (3/55)CM, PCRssu-rRNAC. canis (2), Cryptosporidium spp. (1)De Lucio et al. [81]
Norway1 (1/170)CM, PCRssu-rRNAC. canis (1)Myšková et al. [91]
Domestic ungulates
Giardia duodenalisBos taurusThe Netherlands9 (57/628)CM, PCRgdhA (5), B (4)NAHuetink et al. [92]
Italy30 (3/10)CM, PCRssu-rRNAE (3)Berrilli et al. [54]
Italy100 (24/24)CM, PCRssu-rRNA, gdh, bgA (12), B (5), E (3), A + B (2), A + E (2)A13 (8), A23 (1), A33 (2), A43 (1), B33 (1), B53 (2), B63 (2), E23 (1), E33 (1), E1 + E23 (1)Lalle et al. [55]
Italy32 (162/503)CM, PCRbg, tpiA (4), E (7), A + E (3)NAGeurden et al. [93]
Spain27 (101/379)CM, PCRgdh, bgE (4)Castro-Hermida et al. [94]
Spain19 (68/362)PCRssu-rRNA, gdh, bgE (15), F (4)Cardona et al. [95]
Spain30 (192/649)CM, PCRbgA (18), E (32)AI (39)Castro-Hermida et al. [96]
Denmark35 (401/1150)CM, PCRssu-rRNA, gdhA (8), E (137)NALangkjær et al. [97]
Belgium31 (259/832)CM, PCRbg, tpiA (43), E (77)A23 (ND), A33 (ND), E23 (ND), E33 (ND)Geurden et al. [98]
France40 (190/477)CM, PCRbg, tpiA (8), E (4), A + E (8)NAGeurden et al. [93]
Germany51 (274/536)CM, PCRbg, tpiA (8), E (9), A + E (7)NAGeurden et al. [93]
Germany73 (110/152)PCRssu-rRNA, gdh, bgA (8), E (101), A + E (1)A13 (1), E23 (1), E33 (8)Gillhuber et al. [99]
United Kingdom55 (305/556)CM, PCRbg, tpiA (3), E (2), A + E (4)NAGeurden et al. [93]
Poland17 (16/86)CM, PCRbgA (1), E (3)NAStojecki et al. [77]
Austria27 (48/177)CM, PCRssu-rRNA, bg, tpiA (1), E (30)NALichtmannsperger et al. [100]
Portugal9 (42/467)CM, PCRgdh, bgA (2), B (1), E (11)A23 (2)Mendonça et al. [101]
Portugal15 (13/87)PCRssu-rRNA, gdh, bg, tpiE (1)This study
Capra hircusThe Netherlands100 (1/1)PCR1ssu-rRNA, gdhE (1)van der Giessen et al. [42]
Spain20 (23/116)CM, PCRgdh, bgE (1)Castro-Hermida et al. [94]
Spain42 (133/315)CM, PCRbg, tpiE (31)Ruiz et al. [102]
Belgium36 (53/148)CM, PCRbg, tpiA (6), E (17), E + A (5)A23 (6), E23 (6), E33 (16)Geurden et al. [103]
Greece40 (103/255)CM, PCRbg, tpiA (1), E (26), A + E (3)NATzanidakis et al. [104]
Equus caballusItaly13 (20/150)CM, PCRssu-rRNAE (20)Veronesi et al. [105]
Italy9 (37/431)PCRssu-rRNA, bgA (16), B (11), E (10)A13 (3), B1-23 (8), B1-63 (2), E33 (1)Traversa et al. [106]
Belgium14 (19/134)CM, PCRbg, tpiA (8), B (3)AI (6), AII (2), BIV (3)Kostopoulou et al. [107]
Germany10 (3/30)CM, PCRbg, tpiNANAKostopoulou et al. [107]
Greece12 (22/190)CM, PCRbg, tpiA (4), B (9), E (2)AI (3), AII (1), BIV (9)Kostopoulou et al. [107]
The Netherlands11 (5/44)CM, PCRbg, tpiA (1), B (1)AII (1)Kostopoulou et al. [107]
Poland10 (1/10)CM, PCRbgE (1)Stojecki et al. [77]
Portugal8 (2/26)PCRssu-rRNA, gdh, bg, tpiE (1)This study
Ovis ariesItaly2 (5/325)CM, PCRgdh, bgA (5)AI (5)Giangaspero et al. [108]
Italy100 (3/3)PCR1tpiB (3)B12Aloiso et al. [10]
The Netherlands100 (2/2)PCR1ssu-rRNA, gdhE (2)van der Giessen et al. [42]
Spain33 (118/575)CM, PCRbgE (NA)Castro-Hermida et al. [109]
Spain19 (86/446)CM, PCRgdh, bgE (11), B (1)NACastro-Hermida et al. [94]
Spain42 (162/386)CM, PCRbgA (1), E (74)A13 (1), E13 (2), E23 (2)Gómez-Muñoz et al. [110]
Spain30 (112/377)CM, PCRbgB (11), E (20)Castro-Hermida et al. [96]
Spain89 (107/120)CM, PCRssu-rRNA, gdh, bg, tpiA (59), E (21), A + E (27)AI (7), AII (1)Gómez-Muñoz et al. [111]
Belgium26 (36/137)CM, PCRbg, tpiA (2), E (4), E + A (2)A23 (2), E23 (1), E33 (5)Geurden et al. [103]
Norway22 (244/1095)CM, PCRgdh, bgB (1), E (47)NARobertson et al. [112]
Greece37 (160/429)CM, PCRbg, tpiA (1), E (35), A + E (3)NATzanidakis et al. [104]
Poland22 (18/81)CM, PCRbgA (6), E (10)NAStojecki et al. [77]
Portugal17 (8/46)PCRssu-rRNA, gdh, bg, tpiE (2)This study
Cryptosporidium spp.Bos taurusPoland43 (146/342)CM, PCRssu-rRNAC. muris (1), C. felis (1)Bornay-Llinares et al. [113]
Poland17 (119/700)PCRssu-rRNA, cowp, LIB13C. bovis (52), C. parvum (36), C. andersoni (17), C. ryanae (8)Rzeżutka and Kaupke [114]
Poland10 (76/779)PCRssu-rRNA, cowp, LIB13, gp60C. parvum (76)IIa (64), IId (7), IIl (5)Kaupke and Rzeżutka [115]
Poland45 (725/1601)PCRssu-rRNA, gp60C. parvum (98), C. parvum + C. bovis (2), Cryptosporidium spp. (625)IIa (80), IId (2)Kaupke and Rzeżutka [116]
Czech Republic100 (4/4)PCR1ssu-rRNA, ITS1, hsp70C. muris (4)Morgan et al. [117]
Czech Republic100 (11/11)PCR1ssu-rRNA, hsp70C. andersoni (9), C. parvum, (2)Ryan et al. [118]
Czech Republic100 (2/2)PCR1ssu-rRNA, cowpC. parvum (1), C. andersoni (1)Hajdušek et al. [46]
Czech Republic5 (49/995)CM, PCRssu-rRNA, gp60C. andersoni (41), C. bovis (2), C. parvum (1)IIa (1)Ondráčková et al. [119]
Czech Republic21 (159/750)CM, PCRssu-rRNA, gp60C. parvum (137), C. andersoni (21), C. bovis (3)IIa (131)Kváč et al. [120]
The Netherlands9 (54/628)CM, PCRcowpC. parvum (27)Huetink et al. [92]
The Netherlands100 (160/160)PCR1ssu-rRNA, hsp70, cowp, gp60C. parvum (160)IIa (129), IIj (1)Wielinga et al. [121]
The Netherlands17 (69/399)PCRssu-rRNA, gp60C. parvum (59), C. bovis (6), C. ryanae (4)IIa (47)Pinto et al. [122]
Portugal23 (129/553)CM, PCRcowp, TRAP-C1C. parvum (14)Fonseca et al. [123]
Portugal100 (35/35)PCR1ssu-rRNA, gp60C. parvum (35)IIa (29), IId (3)Alves et al. [124]
Portugal98 (40/41)PCR1gp60C. parvum (40)IIa (39), IId (1)Alves et al. [125]
Portugal18 (82/467)CM, PCRssu-rRNA, hsp70C parvum (78), C. meleagridis (1), C. andersoni (1)Mendonça et al. [101]
Denmark31 (90/292)CM, PCRssu-rRNAC. andersoni (59), C. parvum (21), C. andersoni + C. parvum (10)Enemark et al. [126]
Denmark56 (108/193)CM, PCRcowp, ssu-rRNAC. parvum genotype II (108)Enemark et al. [127]
Denmark29 (336/1150)CM, PCRssu-rRNA, hsp70C. parvum (79), C. bovis (57), C. deer-like genotype4 (11), C. suis (2)Langkjær et al. [97]
Ireland7 (21/288)CM, PCRssu-rRNAC. andersoni (11), C. parvum genotype II (10)Moriarty et al. [128]
Ireland37 (291/779)CM, PCRssu-rRNA, gp60C. parvum (215), C. bovis (5), C. deer-like genotype4 (3)IIa (215)Thompson et al. [129]
Ireland25 (342/1368)CM, PCRssu-rRNA, gp60C. parvum (84), C. bovis (4), C. ryanae (1)IIa (265)De Waele et al. [130]
Ireland66 (68/103)PCR1ssu-rRNA, gp60C. ryanae (36), C. bovis (23), C. parvum (16), C. andersoni (4), Cryptosporidium pig genotype5 (1), Cryptosporidium spp. (19), C. bovis + C. xiaoi (17), C. parvum + C. hominis (3), C. ryanae + C. bovis (2)IIa (9)Mirhashemi et al. [131]
United Kingdom16 (16/101)CM, PCRssu-rRNAC. andersoni (7)Robinson et al. [132]
Serbia/Montenegro60 (62/103)PCRssu-rRNA, cowp, gp60C. parvum (62)IIa (38), IIj (21), IId (3)Misic and Abe [133]
Spain14 (41/291)CM, PCRhsp70C. parvum (41)Castro-Hermida et al. [134]
Spain8 (32/379)CM, PCRssu-rRNA, hsp70C. parvum (10)Castro-Hermida et al. [94]
Spain58 (166/287)CM, PCRssu-rRNA, gp60C. parvum (147), C. bovis (2)IIa (138), IId (2)Quilez et al. [135]
Spain49 (30/61)CM, PCRssu-rRNA, gp60C. parvum (27)IIa (27)Díaz et al. [136]
Spain15 (97/649)CM, PCRssu-rRNAC. parvum (41) C. andersoni (17)Castro-Hermida et al. [96]
Spain94 (131/140)PCR1ssu-rRNA, gp60C. parvum (131)IIa (129), IId (2)Quílez et al. [137]
Spain12 (45/362)PCRssu-rRNAC. felis (4), C. bovis (2), Cryptosporidium spp. (3)Cardona et al. [95]
Spain17 (99/594)PCRssu-rRNA, gp60C. parvum (42), C. bovis (36), C. ryanae (10), C. occultus (7), C. andersoni (2), C. xiaoi (1)IIa (30)Díaz et al. [138]
Belgium19 (155/832)CM, PCRssu-rRNA, hsp70C. parvum (105), C. bovis (9), C. suis (1)IIa (89), IId (1)Geurden et al. [139]
Belgium18 (63/355)PCRssu-rRNA, gp60C. parvum (45), C. bovis (14), C. ryanae (4)IIa (39)Pinto et al. [122]
Hungary49 (39/79)CM, PCRssu-rRNA, gp60C. parvum (21), Cryptosporidium deer-like genotype4 (1)IIa (19), IId (2)Plutzer et al. [140]
Germany36 (48/134)CM, PCRssu-rRNA, cowp, gp60C. parvum (53)IIa (52), IId (1)Broglia et al. [141]
Germany89 (455/512)CM, PCRssu-rRNA, gp60C. parvum (395)IIa (395)Holzhausen et al. [142]
Germany87 (233/268)PCR1ssu-rRNA, gp60C. parvum (233)IIa (226)Göhring et al. [143]
SloveniaND (51/ND)PCRssu-rRNA, gp60C. parvum (45), C. bovis (3), C. ryanaeIIa (41), IIl (4)Soba and Logar [144]
Italy8 (155/2024)CM, PCRcowp, gp60C. parvum (101)IIa (62)Duranti et al. [145]
Italy100 (122/122)PCR1cowpC. parvum (122)Drumo et al. [146]
Italy39 (57/147)CM, PCRssu-rRNA, gp60C. parvum (46), C. bovis (2)IIa (33), IId (3)Díaz et al. [147]
France35 (147/422)PCRssu-rRNA, gp60C. parvum (60), C. ryanae (39), C. bovis (37), C. ubiquitum (1)IIa (51)Follet et al. [148]
France46 (84/182)CM, PCRssu-rRNA, gp60C. ryanae (22), C. bovis (15), C. parvum (14), C. bovis + C. ryanae (5), C. bovis + C. parvum (4), C. parvum + C. ryanae (1)IIa (15)Rieux et al. [149]
France31 (92/300)CM, PCRssu-rRNA, gp60C. parvum (80), C. bovis (2)IIa (52)Rieux et al. [150]
France36 (32/88)CM, PCRssu-rRNAC. bovis (27), C. ryanae (4), C. parvum (1)Rieux et al. [151]
France64 (201/312)CM, PCRssu-rRNA, gp60C. parvum (80), C. bovis (53), C. ryanae (19), C. bovis + C. parvum (4), C. bovis + C. ryanae (4)IIa (51)Rieux et al. [152]
France21 (86/412)CM, PCRssu-rRNA, gp60C. parvum (71), C. hominis (15)IIa (71), Ib (15)Razakandrainibe et al. [153]
France89 (31/35)CM, PCRssu-rRNA, gp60C. parvum (30)IIa (24), IId (3)Mammeri et al. [154]
France23 (79/350)PCRssu-rRNA, gp60C. parvum (51), C. bovis (22), C. ryanae (4), C. andersoni (1), C. xiaoi (1)IIa (47)Pinto et al. [122]
Romania25 (65/258)CM, PCRssu-rRNA, gp60C. parvum (65)IIa (13)Imre et al. [155]
Romania59 (17/29)PCR1ssu-rRNA, gp60C. parvum (17)IIa (1), IId (16)Vieira et al. [156]
Sweden63 (110/176)PCR1ssu-rRNA, gp60C. bovis (83), C. parvum 15), C. ryanae (10), C. andersoni (2)IIa (7), IId (6)Silverlås et al. [157]
Sweden31 (242/782)CM, PCRssu-rRNA, gp60C. parvum (171), C. bovis (7), C. bovis+ C. parvum (5)IIa (136), IId (40)Silverlås et al. [158]
SwedenND (ND/480)CM, PCRssu-rRNA, gp60C. bovis (48), C. ryanae (11), C. parvum (2)IIa (2)Silverlås and Blanco-Penedo [159]
Sweden37 (122/332)CM, PCRssu-rRNA, gp60C. bovis (72), C. ryanae (13), C. parvum (8), C. ubiquitum (1), C. bovis + C. parvum (13), C. ryanae + C. parvum (6)IIa (7), IId (16)Björkman et al. [160]
Sweden39 (92/238)CM, PCRssu-rRNA, gp60C. bovis (63), C. ryanae (7), C. bovis + C. ryanae (2)NAÅberg et al. [161]
Sweden22 (99/455)CM, PCRssu-rRNA, gp60C. bovis (46), C. ryanae (7), C. bovis + C. ryanae (2)NAÅberg et al. [162]
Estonia18 (9/49)CM, PCRssu-rRNA, hsp70, gp60C. parvum (1)IIa (1)Lassen et al. [163]
Estonia23 (110/486)PCRssu-rRNA, gp60C. parvum (105), C. bovis (4), C. ryanae (1)IIa (90), IIl (5)Santoro et al. [164]
Slovakia14 (14/100)PCRssu-rRNA, gp60C. parvum (10), C. bovis (4)IIa (10)Danišová et al. [165]
Austria55 (98/177)CM, PCRgp60C. parvum (37)IIa (35)Lichtmannsperger et al. [100]
Austria55 (98/177)CM, PCRssu-rRNA, gp60C. parvum (67), C. ryanae (11), C. bovis (7)IIa (65)Lichtmannsperger et al. [166]
Cyprus44 (106/242)PCRssu-rRNA, gp60C. parvum (50), C. bovis (23), C. ryanae (28), C. ryanae + C. parvum (5)IIa (47)Hoque et al. [167]
Latvia34 (313/926)CM, PCRssu-rRNA, gp60C. parvum (62), C. bovis (29), C. andersoni (22), C. ryanae (11), C. scrofarum (1), C. ubiquitum (1), C. parvum + C. bovis (3), C. bovis + C. ryanae (3), C. parvum + C. andersoni (1), C. parvum + C. ryanae (1), C. bovis + C. andersoni (1)IIa (55), IId (3)Deksne et al. [168]
Capra hircusCzech Republic100 (1/1)PCR1ssu-rRNA, cowpC. parvum (1)Hajdušek et al. [46]
Spain40 (2/5)CM, PCRssu-rRNAC. xiaoi (2)Díaz et al. [169]
Spain100 (17/17)PCR1ssu-rRNA, gp60C. parvum (17)IId (17)Quílez et al. [135]
Spain63 (74/118)CM, PCRssu-rRNA, gp60C. parvum (60), C. xiaoi (4), C. parvum + C. xiaoi (1)IIa (55), IId (3)Díaz et al. [170]
Spain6 (14/234)PCRssu-rRNA, gp60C. parvum (3), C. ubiquitum (5), C. xiaoi (5)IIa (1), IId (2), XIIa (3)Díaz et al. [171]
Belgium10 (14/148)CM, PCRssu-rRNA, hsp70, gp60C. parvum (11)IIa (3), IId (8)Geurden et al. [103]
The Netherlands100 (1/1)PCR1hsp70, gp60, ML1C. parvum (1)IIa (1)Wielinga et al. [121]
Italy100 (21/21)PCR1cowpC. parvum (21)Drumo et al. [146]
Norway21 (4/19)CM, PCRssu-rRNA, LIB13, gp60C. parvum (3), C. xiaoi (1)IIa (3)Lange et al. [172]
France24 (61/254)CM, PCRssu-rRNA, gp60C. xiaoi (18), C. parvum (1)NARieux et al. [173]
FranceNA (22/ND)CM, PCRssu-rRNAC. ubiquitum (12)Paraud et al. [174]
Greece7 (18/255)CM, PCRssu-rRNA, hsp70, gp60C. xiaoi (7), C. ubiquitum (5), C. parvum (2)IId (2)Tzanidakis et al. [104]
Greece28 (41/148)CM, PCRssu-rRNA, gp60C. parvum (16), C. xiaoi (1)IIa (5), IId (11)Papanikolopoulou et al. [175]
Poland37 (39/105)PCRssu-rRNA, gp60C. xiaoi (29), C. parvum (1)IId (1)Kaupke et al. [176]
Equus caballusCzech Republic100 (3/3)PCR1ssu-rRNA, cowpC. parvum (3)[46]
Czech Republic/Poland3 (12/352)PCRssu-rRNA, gp60C. muris (9), C. parvum (1), C. tyzzeri (1), Cryptosporidium horse genotype (1)IIa (1), IXb (1), VIa (1)Wagnerová et al. [177]
Italy8 (12/540)CM, PCRcowpC. parvum (12)Veronesi et al. [105]
Italy2 (4/185)CM, PCRcowpC. parvum genotype II (4)Perrucci et al. [178]
Italy38 (14/37)PCRssu-rRNA, gp60Cryptosporidium horse genotype (11), C. parvum (3)VIa (9)Caffara et al. [179]
Italy17 (35/205)CM, PCRssu-rRNA, gp60C. parvum (5), Cryptosporidium. horse genotype (21), C. parvum + C. horse genotype (9)IIa (6), IId (2), VIa (20)Galuppi et al. [180]
Belgium5 (6/134)CM, PCRssu-rRNA, hsp70Cryptosporidium horse genotype (6)Kostopoulou et al. [107]
Greece1 (2/190)CM, PCRssu-rRNA, hsp70Cryptosporidium horse genotype (2)Kostopoulou et al. [107]
Ireland56 (23/41)PCR1ssu-rRNA, gp60C. ryanae (17), C. parvum (4), C. bovis (1), Cryptosporidium horse genotype (1), C. pig genotype6 (1), C. bovis + C. xiaoi (5), C. andersoni + C. bovis (1), C. andersoni + C. bovis (1)IIj (1)Mirhashemi et al. [131]
Portugal21 (3/14)CM, PCRssu-rRNA, hsp70C. parvum (1)IIa (1)Couso-Pérez et al. [181]
Spain11 (7/65)CM, PCRssu-rRNA, hsp70C. parvum (6)IIa (2)Couso-Pérez et al. [181]
Ovis ariesPoland10 (16/159)CM, PCRssu-rRNAC. spp. (10)Majewska et al. [182]
Poland19 (45/234)PCRssu-rRNA, gp60C. xiaoi (30), C. bovis (9), C. ubiquitum (3), C. xiaoi + C. parvum + C. hominis (1), C. xiaoi + C. parvum (1), C. xiaoi + Cryptosporidium spp. (1)IIa (2)Kaupke et al. [176]
Denmark100 (1/1)PCR1cowp, ssu-rRNAC. parvum (1)Enemark et al. [127]
Portugal100 (2/2)PCR1ssu-rRNA, gp60C. parvum (2)IIa (1), IId (1)Alves et al. [125]
Belgium13 (18/137)CM, PCRssu-rRNA, hsp70, gp60Cryptosporidium cervine genotype7 (9), C. parvum (1)IIa (1)Geurden et al. [103]
United Kingdom43 (127/297)CM, PCRssu-rRNA, cowpC. parvum (52), C. bovis (5), Cryptosporidium cervine genotype7 (1)Mueller-Doblies et al. [183]
Italy17 (26/149)CM, PCRcowpC. parvum (26)Paoletti et al. [184]
Italy100 (21/21)PCR1cowpC. parvum (21)Drumo et al. [146]
Italy4 (1/27)CM, PCRgp60C. parvum (4)IIa (4)Cacciò et al. [185]
Italy10 (92/915)CM, PCRssu-rRNA, gp60C. parvum (11), C. ubiquitum (4)IIa (5), IId (1), XIIa (4)Dessì et al. [186]
The Netherlands100 (1/1)PCR1hsp70, gp60, ML1C. parvum (1)IIa (1)Wielinga et al. [121]
Spain100 (131/131)PCR1ssu-rRNA, gp60C. parvum (131)IIa (3), IId (128)Quílez et al. [187]
Spain31 (39/127)CM, PCRssu-rRNA, gp60C. parvum (14), Cryptosporidium cervine genotype7 (9)IIa (12)Díaz et al. [136]
Spain19 (42/227)CM, PCRssu-rRNAC. parvum (27)Castro-Hermida et al. [96]
Spain32 (54/171)CM, PCRssu-rRNA, gp60C. parvum (31), C. ubiquitum (10), C. parvum + C. ubiquitum (1)IIa (26)Díaz et al. [170]
Spain6 (19/324)PCRssu-rRNA, gp60C. parvum (13), C. ubiquitum (1), C. xiaoi (3)IIa (8), XIIa (1)Díaz et al. [171]
Norway15 (160/1095)CM, PCRssu-rRNA, actinCryptosporidium cervine genotype7 (35), C. xiaoi (7)Robertson et al. [112]
Norway100 (2/2)CM, PCRssu-rRNA, LIB13, gp60C. parvum (1), C. xiaoi (1)IIa (2)Lange et al. [172]
Romania14 (24/175)CM, PCRssu-rRNA, gp60C. parvum (20), C. ubiquitum (2), C. xiaoi + C. bovis (2)IIa (3), IId (4)Imre et al. [188]
Greece5 (22/429)CM, PCRssu-rRNA, hsp70, gp60C. parvum (7), C. ubiquitum (3), C. parvum (7)IId (7)Tzanidakis et al. [104]
Greece30 (39/132)CM, PCRssu-rRNA, gp60C. parvum (16)IIa (9), IId (4)Papanikolopoulou et al. [175]
Ireland49 (51/104)PCR1ssu-rRNA, gp60C. parvum (14), C. xiaoi (10), C. ryanae (9), C. ubiquitum (7), C. bovis (1), Cryptosporidium spp. (15), C. bovis + xiaoi (18)IIa (6)Mirhashemi et al. [131]
France100 (23/23)CM, PCRssu-rRNA, gp60C. parvum (23)IIa (23)Mammeri et al. [189]
France35 (53/151)CM, PCRssu-rRNA, LIB13, gp60C. parvum (10), C. xiaoi (4), C. ubiquitum (1)IIa (3), IId (2)Bordes et al. [190]

Note. bg, β-giardin; CM, conventional microscopy; cowp, Cryptosporidium oocyst wall protein; gdh, glutamate dehydrogenase; gp60, 60-kDa glycoprotein; hsp70, 70-kDa heat shock protein; ITS, internal transcribed spacer; LIB13, protein of unknown function; NA, no amplification; ND, no data available; PCR, polymerase chain reaction; ssu-rRNA, small subunit ribosomal RNA; TRAP-C1, thrombospondin-related adhesive protein of Cryptosporidium-1; tpi, triose phosphate isomerase. 1PCR conducted only in samples with a previous positive result by another detection method. 2Primer sequences specifically designed to amplify assemblage D. 3Previous nomenclature adopted by the authors. 4Current C. ryanae. 5Current C. suis. 6Current C. scrofarum. 7Current C. ubiquitum.