Catch-All No More: Integrative Systematic Revision of the Genus Allolobophora Eisen, 1874 (Crassiclitellata, Lumbricidae) with the Description of Two New Relict Earthworm Genera

Table 3

Comparison of the main morphological characters of the four revised genera and its closest genera.

	Allolobophora Eisen, 1874	Gatesona Qiu and Bouché, 1998	Eophila Rosa, 1893	Cernosvitovia Omodeo, 1856	Panoniona Mršić and Šapkarev, 1988	Vosgesia Martínez Navarro and Marchán, 2022	Avelona Qiu and Bouché, 2000	Heraultia Martínez Navarro and Marchán, 2022	Prosellodrilus Bouché, 1972	Cataladrilus Qiu and Bouché, 1998
Length (mm)	30-150	68-192	100-360	42-250	40-151	50-75	40-80	50-75	35-335	25-220
N. segments	60-210	118-197	139-341	138-304	115-181	125-140	100-125	156-172	130-350	124-384
Pigmentation	Green (sometimes absent)	Pigmentary dots	Purplish-brown in bands or brown	Light grey to brown	Absent	Absent	Brown	Absent	Absent	Absent
Prostomium	Epilobous	Epilobous	Epilobous	Epilobous or prolobous	Epilobous	Epilobous	Epilobous	Epilobous	Epilobous, prolobous or tanilobous	Epilobous
First dorsal pore	(3/4), 4/5, (5/6), (6/7)	4/5-7/8	5/6	9/10, 10/11, 12/13, 13/14	4/5	4/5	8/9	4/5	4/5-6/7	4/5-11/12
Spermathecal pores	Simple in 7/8, 8/9, 9/10, 10/11	Simple in 9/10, 10/11	Paired in 9/10, 10/11	Position and number highly variable, from 9/10 to 19/20	Double or multiple in 9/10, 10/11	Simple or double in 9/10, 10/11	Simple in 9/10, 10/11	Simple in 10/11, 11/12, 12/13	Simple in 12/13, 13/14, and 14/15 or simple, double, or multiple in 14, 15	Simple in 9/10, 10/11
Clitellum	28-78	1/2 22-36	24-44	21-65	23-34	26-34	29-36	21-36	(19) 20-27 (34)	22-46
Tubercula pubertatis	30-78	27-26	29-40	21-62	28-34	27-33	(31) 32-35 (36)	27-29	Variable	28-45
Oesophageal hearts	6-11	5, 6-11, 12	6-11	5, 6-11, 12	6-11	6-11	6-11	6-11	6, 7-11	6, 7-11
Calciferous glands	10-14 with diverticula in 10	10-12 (14) with diverticula in 10	10-12 with diverticula in 10	10-13	10-14 with diverticula in 10	10-14 with diverticula in 10	10-14 with diverticula in 10	13-14	11-15 with diverticula in 11	11-15 with diverticula in 11
Crop	15-16	15-16	15-16	15-16	15-16	15-16	15-16	15-16	15-16	15-16 (17, 18)
Gizzard	17-18	17-18	17-19	17-19, 20	17-18	17-18	17-18	(17) 18	17-18	17 (18, 19)-18 (21)
Typhlosole	Bifid or trifid	Bifid	Trifid	Trifid	Trifid	Bifid	Bifid	Bifid	Bifid or multifid	Simple, bifid, trifid, or multifid
Seminal vesicles	9-12	(9), (10), 11, 12	9-12	9–12 or 11–12	11-12	11-12	11-12	11-12	11-12	11-12
Spermathecae	Simple in 7/8, 8/9, 9/10, 10/11	Simple, globular in 9, 10	Simple in 9/10, 10/11	Position and number highly variable, from 9/10 to 19/20.	Double or multiple in 10, 11	Simple or double in 10, 11	Simple in 10, 11	Globular in 11, 12, 13	Variable in (12/13), 13/14, 14/15	9/10, 10/11