Research Article

Catch-All No More: Integrative Systematic Revision of the Genus Allolobophora Eisen, 1874 (Crassiclitellata, Lumbricidae) with the Description of Two New Relict Earthworm Genera

Table 4

List of the species included in Allolobophora sensu stricto and its main morphological features.

All. chloroticaAll. burgondiaeAll. ictericaAll. molleriAll. moebiiAll. dubiosa

Length (mm)30-8075-8070-90150-20060-125126-240
N. segments60-140150-170140-200145-210104-122121-303
PigmentationGreen (sometimes absent)Absent/brownishAbsentGreenGreenAbsent
ProstomiumEpilobousEpilobousEpilobousEpilobousEpilobousEpilobous
First dorsal pore4/5 (5/6)4/54/54/5 (5/6)4/5 (5/6)4/5
Spermathecal pores8/9, 9/10, 10/118/9, 9/10, 10/11(7/8), 8/9, 9/10, 10/117/8, 8/9, 9/10, 10/117/8, 8/9, 9/10, 10/11(7/8), 8/9, 9/10, 10/11
Clitellum(28) 29-37(27) 28-38 (39)(33) 34-43 (44)(48) 49-59(52) 53-61 (62)(36) 38-46 (49)
Tubercula pubertatis30-3630-37(35) 36-42 (43)50-57(55) 56-61(43) 44-47 (48)
Oesophageal hearts6-116-116-116-116-116-11
Calciferous glands10-14 with diverticula in 1010-14 with diverticula in 1010-14 with diverticula in 1010-12 with diverticula in 1010-12 with diverticula in 1010-14 with diverticula in 10
Crop15-1615-1615-1615-1615-1615-16
Gizzard17-1817-1817-1817-1817-1817-18
TyphlosoleBifidTrifidBifidTrifidTrifidBifid
Nephridial bladdersU-shapedU-shapedU-shapedU-shapedU-shapedU-shaped
Seminal vesicles9, 10, 11, 129, 10, 11, 129, 10, 11, 129, 10, 11, 129, 10, 11, 129,10, 11, 12
Spermathecae9, 10, 119, 10, 119, 10, 117/8, 8/9, 9/10, 10/117/8, 8/9, 9/10, 10/118/9, 9/10, 10/11